Educated at Hymers Collge Hull, before stuying medicine and biochemistry at the University of Edinburgh.

ACADEMIC AND PROFESSIONAL HISTORY

1971-72       Surgical and Medical House Officer, Longmore and Western General Hospitals, Edinburgh

1972-75       Faculty of Medicine Research Scholar, University of Edinburgh (supervisor Dr I B R Bowman)

1975-76       MRC Travelling Fellow, University of Amsterdam (Prof Piet Borst)

1976-80       Research Fellow, University of Edinburgh (Dr I B R Bowman)

1980-81       Research Fellow, London School of Hygiene and Tropical Medicine (Prof W Peters)

1981-87       Assistant Professor, The Rockefeller University, New York (Prof A Cerami)

1987-90       Senior Clinical Lecturer, London School of Hygiene & Tropical Medicine

1990-96        Professor of Molecular Parasitology and Head, Biochemistry & Chemotherapy Unit, LSHTM

1996-           Wellcome Principal Research Fellow, Professor of Biochemistry and Head of Division of Biological Chemistry & Drug Discovery (1996-2102), University of Dundee

2006-           Co-Director Drug Discovery Unit, University of Dundee

HONOURS AND AWARDS

1987       Seymour H. Hutner Prize, Society of Protozoologists

1988       Frederick Murgatroyd Prize, Royal College of Physicians

1989       Fellow of the Linnean Society, London

1991       Ernst A.H. Friedheim Commemorative Lecture, Rockefeller University, New York

1993       Co-vice chairman (with Immo Scheffler) Gordon Research Conference on Polyamines

1995       Co-chairman (with Immo Scheffler) Gordon Research Conference on Polyamines

1996       John Scaife Commemorative Lecture, Edinburgh

1996       C.A.Wright Medal, British Society for Parasitology

1999       Foreign Corresponding Member, National Academy of Medicine of Buenos Aires, Argentina

2001       Fellow of the Royal Society of Edinburgh

2005       Fellow of the Academy of Medical Sciences

2005       Kitasato Microbial Chemistry Medal, Kitasato Institute, Japan

2005       Commander of the British Empire for services to medical science

2006       3Rs Prize 2006 for Refinement and Reduction of Animal Models of Leishmaniasis

2011       Fellow of the Society of Biology