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The Strange Case of the Noncanonical Lamina: Deep Divisions in Nuclear Organisation?

Research output: Chapter in Book/Report/Conference proceedingChapter

Abstract

The nuclear envelope is subtended in most eukaryotes by a proteinaceous lamina, a network that has been recognised since the 1950s. Originally considered as a simple structural support, it is now clear that the lamina can be highly dynamic and participates in a multitude of functions, including transcriptional and epigenetic regulation, definition of chromatin domains, genome stability and the positioning of nuclear pore complexes. The major protein components of the lamina in metazoans are ~60 kDa lamins, which assemble to form fibres and a network and are regulated by multiple mechanisms. Despite a broad taxonomic distribution beyond Metazoa, lamins are not universal and, in at least three major lineages, are absent, specifically fungi, plants and kinetoplastid protists. The latter two possess lineage-specific lamin analogues, the CRWN and NUP-1/NUP-2 proteins, respectively. Here we discuss and compare the kinetoplastid and plant lamina, their origins, components and functions and spectacular examples of convergent evolution.

Original languageEnglish
Title of host publicationLamins in Genome Organizatioin and Diseases
EditorsKaushik Sengupta, Antoine Muchir
PublisherSpringer Science and Business Media B.V.
Chapter8
Pages185-200
Number of pages16
Volume115
ISBN (Electronic)9783032005373
ISBN (Print)9783032005366
DOIs
Publication statusE-pub ahead of print - 1 Oct 2025

Publication series

NameSubcellular Biochemistry
Volume115
ISSN (Print)0306-0225

Keywords

  • CRWN proteins
  • Eukaryogenesis
  • Evolution
  • Gene expression
  • Lamina
  • Nuclear pore complex
  • Nucleus
  • Plants
  • Trypanosomes

ASJC Scopus subject areas

  • Biochemistry
  • Cell Biology

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